Y-DNA Haplogroup D and its Subclades - 2016

The entire work is identified by the Version Number and date given on the Main Page. Directions for citing the document are given at the bottom of the Main Page.

Version History     Last revision date for this specific page: 24 May 2016

Because of continuing research, the structure of the Y-DNA Haplogroup Tree changes and ISOGG does its best to keep the tree updated with the latest developments in the field. The viewer may observe other versions of the tree on the Web. Email Ray Banks if the differences need clarification or if you find any broken links on this page.

LINKS:  Main Page   Y-DNA Tree Trunk    SNP Index    Papers/Presentations Cited    Glossary   Listing Criteria
SNP SYMBOLS:  Not on 2015 tree  Confirmed within subclade  Investigational items 

^ Indicates a next-generation sequencing entry which does not yet meet quality guidelines for minimum number of reads.
^^ Indicates an entry which does not meet quality guidelines but may be helpful.
~ Indicates only an approximate location on the tree.

The criteria for a representative SNP printed in bold for a subclade is: traditional usage, testing one in multiple labs, and/or being found in the area of the chromosome used in recent research studies.

SNPs listed below in italics (colored black or red) are quality variants from next-generation sequencing reports consistently showing as representing that subgroup.

Contact Person for Haplogroup D: Ray Banks

Link to Experimental Composite Y-DNA Haplogroup D Tree by Ray Banks.

D   M174/Page30, A743.2/FGC1224.2, CTS94, CTS1014.2, CTS1171, CTS1269, CTS1453, CTS1869, CTS2671, CTS2682^, CTS2741, CTS2985, CTS3942/Z1604, CTS4030/V2895/Z1605, CTS4217, CTS5001, CTS5085^/Z1607^, CTS5704, CTS5725^, CTS7082, CTS7660/F3709, CTS9290, CTS9434^, CTS10128,  CTS11261, F887, F975, F1034/V1320, F1066/V1437, F1137/V1774/Z1591, F1163/Z1592, F1264/V2153, F1278/V2177, F1306/V2256, F1344/V2338, F1379/V2446, F1439, F1447, F1533, F1612, F1620.1, F1539, F1540, F1548/Z1594, F1642, F1919/V2849, F1927/V2885, F2062, F2147/Z1608, F2157, F2161/Z1610, F2285, F2318, F2325, F2455, F2473, F2662/Z1612, F2765, F2831/V3465, F2923/V3694, F2956, F3289, F3321, F3328/Z1620, F3370, F3519, F3615, F3666, F3835, F3845/Z1588, F3920, F4030, FGC6233/Y11929,  FGC6254/V1375/Y11253, FGC6260/Y12042, FGC6275/Y11254, FGC6292/Y11256, IMS-JST021355, V1288/Z3711,  V1852/Z3720,  Z1590, Z1618, Z1619, Z3703, Z3704, Z3705, Z3707, Z3715, Z3727, Z3728^, Z3731^, Z3760, Z3762, Z3765, Z3767, Z3771, Z3787, Z3794, Z3795, Z3796, Z17157
D~ CTS260/Z1580, CTS3946, CTS8801, CTS9128, CTS9564, CTS10169, CTS11608, CTS11804, CTS12500, F1690, F3704, Z3701, Z3718, Z3733, Z3772, Z3806
 D1   CTS11577, CTS9290, F2285
  D1a   F6251/Z27276, Z27283
   D1a1   M15
    D1a1a   N1
    D1a1a~ F849, F901, F941, F1182, F1231, F1400, F1709, F1868, F1943, F2025, F2160, F2238, F2460, F2586, F2715, F2762, F3077, F3097, F3306, SK539, Y19636, Y19641, Z27272, Z27274, Z27277, Z27281, Z27282, Z27291, Z27293, Z27295, Z27298, Z27299,  Z27301, Z27305, Z27306, Z27307, Z27308, Z27309, Z27311, Z27312, Z27316, Z27320, Z27321, Z27323, Z27324, Z27335, Z27341, Z27344, Z27346, Z27349, Z27350, Z27351, Z27353, Z29449
     D1a1a1  Z27269, PH4/Z29448, YP4237/Z27338, Z27270, Z27273, Z27275, Z27278, Z27279, Z27280, Z27285, Z27286, Z27287, Z27289, Z27290, Z27292, Z27294, Z27296, Z27297, Z27302, Z27303, Z27304, Z27310, Z27314, Z27315, Z27317, Z27318, Z27319, Z27322, Z27325, Z27326, Z27327, Z27328, Z27329, Z27330, Z27332, Z27333, Z27334, Z27337, Z27339, Z27340, Z27343, Z27345, Z27347, Z27348, Z29450, Z29451, Z29452, Z29453
      D1a1a1a  PH4979, PH344, PH1366, PH2207, PH2452, PH3173, Z29426, Z29427, Z29430, Z29433, Z29436, Z29438, Z29439, Z29441, Z29445, Z29446, Z29447
      D1a1a1a1~ PH1991, Z29428, Z29429, Z29431, Z29432, Z29434, Z29435, Z29437, Z29440
      D1a1a1a1a~ Z31585, Z31586, Z31587
      D1a1a1a1b~ F729, F737, F738, F3828
      D1a1a1a2~ Y7820, Z40573, Z40574, Z40575, Z40576
      D1a1a1b  Z31591, FGC5437.2, SK541, SK2296, YP1043.2, Z31581, Z31582, Z31583, Z31594, Z31595, Z31596, Z31597, Z31598, Z31599, Z31602, Z31603, Z31604, Z31605, Z31606, Z31607, Z31608, Z31609, Z31610, Z31611, Z31612, Z31614, Z31615, Z31616, Z31618, Z31620, Z31621, Z31622, Z31623, Z31627, Z31628
      D1a1a1b1~ Z31625, Z31626
      D1a1a1b2~ Z31584, Z31590, Z31592, Z31593
      D1a1a1c~  Z40603, Z40604, Z40605, Z40606, Z40607 
     D1a1a2  F1070, M5839, Z31624, Z40580, Z40581, Z40582, Z40583, Z40584, Z40585, Z40586, Z40587, Z40588, Z40589, Z40590, Z40591, Z40592, Z40593, Z40594, Z40595, Z40596, Z40597 
      D1a1a2a~ Z40598, Z40599, Z40600, Z40601, Z40602
      D1a1a2b~ CTS5502, CTS6519/S4247, F1531, F3857, Z40608
   D1a2   P99
    D1a2a   P47
     D1a2a1   M533
     D1a2a1~  CTS852.2/PF4504.2, F1287, F1317, F1335, F1412.2, F1771, F1852, F1861.2, F2465, F2835, F2886, F2904, F2967, F3030, M3786, F4339/Y15405/Z34352, PH116/Y14736, PH153, PH232/Y14738, PH396, PH402, PH504/Y14740, PH460, PH648/Y14745, PH824/Y14777, PH893/Y14778, PH906/Y14779, PH937/Y14780, PH1154/Y14783, PH1245, PH1251, PH1281/Y14785, PH1384, PH1394/Y14787, PH1406/Y14788, PH1433/Y14790, PH1451, PH1587, PH1649/Y14797, PH1683, PH1922/Y14805, PH2008/Y14806, PH2062, PH2066, PH2151/Y14807, PH2427, PH2447/Y14810, PH2468/Y14811, PH2607, PH2661, PH2788, PH2795/Y14818, PH2803/Y14819, PH2951/Y14822, PH2974/Y14823, PH3088, PH3166/Y14824, PH3187/Y14825, PH3471, PH3475/Y14829, PH3517, PH3518, PH3654/Y14831, PH3737/Y14833, PH3762/Y14834, PH3831, PH3836/Y14836, PH3858, PH3915/Y14839, PH3936/Y14840, PH3955, PH3998, PH4001/Y14842, PH4246, PH4327, PH4337/Y14847, PH4528/Y14848, PH4533, PH4547/Y14849, PH4717/Y14850, PH4791/Y14852, PH4792, PH4860, PH4866/Y14857, PH4923, PH5000/Y14861, PH5123, PH5131/Y14868, PH5258, PH5290, PH5301, PH5403/Y14876, PH5446, S17259, SK553/Y14732, SK554/Y14761, SK556/Y14760, SK557/Y14759, SK558, SK559/Y14773, SK560/Y14772, SK561, Y14729/Z34142, Y14733/Z34145, Y14734/Z34147, Y14739/Z34156, Y14741/Z34158, Y14742/Z34159, Y14746/Z34161, Y14747/Z34166, Y14748/Z34168, Y14749/Z34169, Y14750/Z34171, Y14751/Z34172, Y14752/Z34174, Y14753/Z34175, Y14754/Z34176, Y14755/Z34177, Y14756/Z34178, Y14757/Z34181, Y14758/Z34184, Y14762/Z34185, Y14763/Z34186, Y14764/Z34187, Y14765/Z34189, Y14766/Z34190, Y14767/Z34191, Y14769/Z34197, Y14770/Z34198, Y14771/Z34201, Y14774/Z34209, Y14775/Z34210, Y14781/Z34221, Y14782/Z34222, Y14789/Z34225, Y14792/Z34228, Y14794/Z34229, Y14795/Z34230, Y14796/Z34232, Y14798/Z34234, Y14800/Z34237, Y14802/Z34242, Y14803/Z34243, Y14804/Z34244, Y14808/Z34247, Y14809/Z34252, Y14813/Z34253, Y14814/Z34255, Y14815/Z34256, Y14816/Z34259, Y14817/Z34260, Y14820/Z34261, Y14821/Z34263, Y14827/Z34268, Y14832/Z34270, Y14837/Z34273, Y14838/Z34274, Y14843/Z34278, Y14844/Z34279, Y14845/Z34280, Y14853/Z34296, Y14854/Z34297, Y14855/Z34301, Y14858/Z34304, Y14859/Z34306, Y14863/Z34308, Y14864/Z34310, Y14865/Z34311, Y14866/Z34314, Y14869/Z34325, Y14870/Z34326, Y14871/Z34327, Y14872/Z34328, Y14873/Z34329, Y14874/Z34331, Y14877/Z34334, Y14878/Z34340, Y14879/Z34341, Y14880/Z34343, Y14882/Z34345, Y14883/Z34347, Y14884/Z34348, Y14885/Z34349, Y14886/Z34354, Y14887/Z34355, Y14888/Z34357, Y15306/Z34309, Y15307/Z34337, Y15309/Z34199, Y15310/Z34200, Y15311/Z34254, Y15312/Z34276, Y15313/Z34277, Y15314/Z34295, Y15315/Z34298, Y15316/Z34299, Y15317/Z34300, Y15318/Z34307, Y15319/Z34335, Y15320/Z34339, Y15321/Z34353, Y15322/Z34275, Y15323/Z34146, Y15324/Z34148, Y15328/Z34195, Y15330/Z34155, Y15331/Z34157, Y15332/Z34162, Y15333/Z34163, Y15334/Z34164, Y15335/Z34165, Y15336/Z34167, Y15338/Z34170, Y15339/Z34173, Y15340/Z34179, Y15341/Z34180, Y15342/Z34182, Y15343/Z34183, Y15344/Z34192, Y15345/Z34193, Y15346/Z34194, Y15347/Z34202, Y15348/Z34204, Y15349/Z34211, Y15350/Z34212, Y15351/Z34219, Y15352/Z34220, Y15355/Z34224, Y15356/Z34226, Y15357/Z34227, Y15359/Z34231, Y15360/Z34233, Y15361/Z34235, Y15362/Z34236, Y15363/Z34238, Y15364/Z34239, Y15365/Z34240, Y15366/Z34246, Y15367/Z34250, Y15368/Z34251, Y15369/Z34257, Y15370/Z34258, Y15371/Z34262, Y15372/Z34264, Y15373/Z34265, Y15375/Z34266, Y15376/Z34267, Y15377/Z34269, Y15378/Z34271, Y15379/Z34272, Y15381/Z34282, Y15385/Z34292, Y15386/Z34293, Y15387/Z34294, Y15389/Z34302, Y15390/Z34303, Y15391/Z34312, Y15392/Z34313, Y15393/Z34315, Y15395/Z34316, Y15398/Z34330, Y15399/Z34332, Y15400/Z34333, Y15401/Z34338, Y15402/Z34342, Y15403/Z34344, Y15404/Z34350, Y15406/Z34356, Y15409/Z34160, Y15412/Z34241, Y15414/Z34305, Y15420/Z34188, Z34217, Z34218, Z34223, Z34248, Z34324, Z34346
      D1a2a1a~  Z34358, Z34359, Z34360, Z34361, Z34362
      D1a2a1b~  PH97, Z34364, Z34365
  D1b   M64.1/Page44.1, CTS69, CTS134/V1058, CTS323,CTS625/Z1587, CTS862, CTS865, CTS1158/Z3706, CTS1827, CTS1828, CTS1841, CTS1877, CTS1904, CTS1910, CTS2061/Z1586, CTS2109, CTS2191/Z1598, CTS2478, CTS2573, CTS2739, CTS2890/Z1600, CTS2960, CTS3177/V2557, CTS3371, CTS3621/V2806, CTS3651, CTS3715^, CTS3823/Z1602, CTS3940/Z1603, CTS4134, CTS4578, CTS4597, CTS4705^, CTS5281^/Z3741^, CTS5828, CTS5829, CTS6051, CTS6157, CTS6220, CTS6242, CTS6248, CTS6277, CTS6332, CTS6448/Z1611, CTS6494, CTS6594, CTS6655, CTS7089, CTS7093, CTS7106, CTS7165, CTS7214, CTS7317,  CTS7508, CTS7626, CTS8060, CTS8210, CTS8337, CTS8348, CTS8435, CTS8505^, CTS8581, CTS8744, CTS8922, CTS9097/Z1614,  CTS9134, CTS9203, CTS9246/V3485, CTS9525, CTS9627, CTS10087^, CTS10197, CTS10502/V4176/Z3820, CTS10703^, CTS10750, CTS10809, CTS10908, CTS10961^, CTS11124, CTS11125, CTS11308,  CTS11834, CTS11915, CTS12007, CTS13016, FGC6245/Y12560, FGC6246/Y12563, FGC6247/Y11725, FGC6251/Y11727, FGC6255/Y11179, FGC6276/Y12545, FGC6278/Y12564, FGC6289/SK543/V1917/Y11720, FGC6299/Y11722, FGC6326/V2498/Y11721, FGC8045/Y12550, L1377, M55, M57, M179/Page31, M359.1/P41.1, M9313^,  P37.1, P190, PF7315/Z1621, V1324/Z3713, V1474/Z3716, V2053/Z1593, V2233/Z3723, Z1589, Z1596, Z1615, Z1616, Z1617, Z3660^,  Z3708, Z3709, Z3710, Z3712, Z3714, Z3722.1/V2156.1, Z3724, Z3725, Z3726^, Z3729, Z3730^, Z3732,  Z3744, Z3749, Z3750, Z3751, Z3752, Z3753, Z3755, Z3756, Z3757, Z3759, Z3761, Z3763, Z3764, Z3766, Z3769, Z3770, Z3797, Z3798, Z3799, Z3783, Z3809
  D1b~  CTS12896, Z3738
   D1b1   M116.1, CTS2728, CTS3978, CTS5304/Z1461, CTS5674/Z1462, CTS5739, CTS6055/Z1463, CTS6238, CTS6485/Z1464, CTS6945, CTS8341, CTS8484, CTS8517, CTS8613, CTS8984, CTS9896/V3805, CTS9988/PF4859/Z1871, CTS10441, CTS10619
    D1b1a   M125
    D1b1a~ IMS-JST75888-GC
      D1b1a1   P42
      D1b1a1a   P12_1, P12_2, P12_3
      D1b1a2   IMS-JST022457, CTS107/IMS-JST055457
      D1b1a2~ Z1476
      D1b1a2a   P53.2
      D1b1a2b   IMS-JST006841/Page3
       D1b1a2b~ IMS-JST119166
       D1b1a2b1   CTS3397, CTS1982/Z1498, CTS3197, CTS4606/Z1499, Z3611, Z3642, Z3643
        D1b1a2b1a   Z1500, CTS8181, CTS9242/Z1502, CTS10268, CTS10511/Z1503, Z1501, Z3603, Z3605, Z3607, Z3610, Z3617, Z3636, Z3651, Z14780, Z14781
         D1b1a2b1a1   Z1504, CTS8093, Z14779
          D1b1a2b1a1a   Z14778
          D1b1a2b1a1a1   FGC6373
         D1b1a2b1a2~ Z40609, Z40610, Z40611, Z40612, Z40613
         D1b1a2b1a3~ Z40614, Z40615, Z40616, Z40617, Z40618
    D1b1b~ M151
    D1b1c~ P120
    D1b1d   CTS6609
     D1b1d1   CTS1897/Z1574, CTS504/Z1569, CTS741, CTS906/Z1570, CTS2296, CTS2798, CTS3097/Z1575, CTS3906, CTS4219, CTS4305, CTS4517, CTS6090, CTS7590/Z1576, CTS8368, CTS8666, CTS10807, Z1568, Z1571, Z1572, Z1573, Z1578, Z1579, Z3838, Z3840, Z3844, Z3851, Z14867, Z14868, Z14869, Z14870, Z14871, Z14872, Z14873, Z14874, Z14875, Z14876, Z14877, Z14878, Z14879, Z14880
      D1b1d1a   CTS218/Z1527, CTS321/Z1528, CTS356/Z1532, CTS621, CTS1228/Z1533, CTS1670/Z1546, CTS2078/Z1547, CTS2378/Z1548, CTS2482, CTS3489/Z1549, CTS3636/Z1550, CTS5307, CTS5862/Z1552, CTS6303/Z1553, CTS6392, CTS6499, CTS6572/Z1554, CTS6859, CTS7234/Z1555, CTS7398, CTS7543, CTS7615/Z1556, CTS7999/Z1557, CTS8230, CTS8960/Z1558, CTS9028, CTS9335, CTS9902, CTS10793/Z1566, CTS11032, CTS11048, CTS11071/Z1567, CTS12720, IMS-JST022456, PF7228, Z1529, Z1530, Z1531, Z1534, Z1535, Z1538, Z1539, Z1540, Z1541, Z1542, Z1560, Z1561, Z1562, Z1563, Z1564, Z1565, Z3834, Z3839, Z3841, Z3843, Z3847, Z3849, Z3850, Z3852, Z3854, Z3855, Z3856, Z14820, Z14821, Z14822, Z14823, Z14824, Z14825, Z14826, Z14827, Z14828, Z14829, Z14830, Z14831, Z14832, Z14833, Z14834, Z14835, Z14836, Z14837, Z14838, Z14839, Z14840, Z14841, Z14842, Z14843, Z14844, Z14845, Z14846, Z14847, Z14848, Z14849, Z14850, Z14851, Z14852, Z14853, Z14854, Z14855, Z14856, Z14857, Z14858, Z14859, Z14860, Z14861, Z14862, Z14863, Z14864, Z14865, Z14866
       D1b1d1a1   CTS4617
        D1b1d1a1a   CTS6909, CTS1614/Z1545, Z14802, Z14804, Z14805, Z14806, Z14807, Z14809, Z14810, Z14811, Z14812, Z14814, Z14818
      D1b1d1b   CTS1964, CTS6511, M6525, Z14881, Z14882, Z14883, Z14884, Z14885, Z14886, Z14887, Z14888
   D1b2  CTS583/Z1516, CTS131, CTS881/Z1517, CTS1352, CTS1592, CTS2098/Z1520, CTS2712, CTS2820, CTS3798, CTS3808, CTS3879, CTS4645/Z1521, CTS6144, CTS6908, CTS7457, CTS7467, CTS8057/Z1523, CTS8075, CTS8327, CTS8635, CTS9143, CTS11368, Z1515, Z1518, Z1519, Z1524, Z1526, Z3801, Z3810, Z3811, Z3822, Z3823, Z3824, Z3827, Z3832, Z3833, Z14793, Z14794, Z14795, Z14796, Z14797, Z14798, Z14799, Z14800, Z17170, Z17171
     D1b2a   CTS220, CTS709, CTS1333, CTS1439/Z1505, CTS2259, CTS2940, CTS3320/Z1508, CTS4158/Z1509, CTS5302, CTS5941, CTS6287/Z1510, CTS6342/Z1511, CTS10054, CTS10126, CTS11474, CTS11619, L495.2/Z2788.2, Z1506, Z1507, Z1514, Z3807, Z3814, Z3819, Z14782, Z14783, Z14784, Z14785, Z14786, Z14787, Z14788, Z14789, Z14790, Z14791, Z14792, Z17172, Z17174, Z17175
     D1b2a1   CTS10495, Z17173, Z17176
     D1b2a2   CTS11285, Z17254, Z17255, Z17256, Z17257, Z17258
  D2  L1366, L1378, M226.2


Y-DNA haplogroup D is seen primarily in Central Asia, Southeast Asia, and in Japan and was established approximately 50,000 years ago. Subgroup D-M15 is seen in Tibet, Mongolia, Central Asia, and Southeast Asia, and the subgroup D-P47 is seen in Central Asia. The subgroup D-M55 is seen almost exclusively in Japan. The high frequency of haplogroup D in Tibet (about 50%) and in Japan (about 35%) implies some early migratory connection between these areas. Examination of the genetic diversity seen in subgroup D-M55 in Japan implies that this group has been isolated in Japan for between 12,000-20,000 years. The highest frequencies of D-M55 in Japan are seen among the Ainu and the Ryukyuans. Subgroup D-L1366 is so far noted only in the Philippines.


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Su et al, Y-chromosome Evidence for a Northward Migration of Modern Humans into Eastern Asia during the Last Ice Age, (pdf), American Journal of Human Genetics, 65:1718-1724, 1999.
Thangaraj et al, Genetic Affinities of the Andaman Islanders, a Vanishing Human Population. (pdf) Current Biology, 13:86-93, 2003.
Xue et al, A Spatial Analysis of Genetic Structure of Human Populations in China Reveals Distinct Difference between Maternal and Paternal Lineages. European Journal of Human Genetics, 16:705-17, 2008.

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ISOGG Wiki - What you need to know about Genetic Genealogy.
D Haplogroup (YDNA) Project, Ray Banks.

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