Y-DNA Haplogroup R and its Subclades - 2011
The entire work is identified by the Version Number and date given on the Main Page.   Directions for citing the document are given at the bottom of the Main Page.
Version History     Last revision date for this specific page: 28 December 2011

Because of continuing research, the structure of the Y-DNA Haplogroup Tree changes and ISOGG does its best to keep the tree updated with the latest developments in the field. The viewer may observe other versions of the tree on the Web. Email Alice Fairhurst if the differences need clarification or if you find any broken links on this page.

Link to Important Announcement of March 21, 2011 on the updating of the majority of subclades in Haplogroup R.


LINKS:  Main Page   Y-DNA Tree Trunk   SNP Index   Papers/Presentations Cited   Glossary   Listing Criteria
CLADE/SUBCLADE SYMBOLS:  Added  Redefined 
SNP SYMBOLS:  Not on 2010 tree  Confirmed within subclade  Provisional  Private  Investigation 

R   M207/Page37/UTY2, P224, P227, P229, P232, P280, P285, S4, S9
•     R*   -
•     R1   M173/P241/Page29, M306/S1, P225, P231, P233, P234, P236, P238, P242, P245, P286, P294
•    •     R1*   -
•    •     R1a   L62/M513, L63/M511, L145/M449, L146/M420
•    •     •      R1a*   -
•    •     •      R1a1   L120/M516, L122/M448, M459, Page65.2/SRY1532.2/SRY10831.2
•    •     •    •     R1a1*   -
•    •     •    •     R1a1a   L168, L449, M17, M198, M512, M514, M515
•    •     •    •     •     R1a1a*   -
•    •     •    •     •     R1a1a1   L457, M417, Page7
•    •     •    •     •    •     R1a1a1*   -
•    •     •    •     •    •     R1a1a1a   M56
•    •     •    •     •    •     R1a1a1b   M157.1
•    •     •    •     •    •     R1a1a1c   M64.2/Page44.2, M87, M204
•    •     •    •     •    •     R1a1a1d   P98
•    •     •    •     •    •     R1a1a1e   PK5
•    •     •    •     •    •     R1a1a1f    M434
•    •     •    •     •    •     R1a1a1g   Z283
•    •     •    •     •    •     •     R1a1a1g*   -
•    •     •    •     •    •     •     R1a1a1g1   M458
•    •     •    •     •    •     •    •     R1a1a1g1*   -
•    •     •    •     •    •     •    •     R1a1a1g1a   M334    (position relative to L260 uncertain)
•    •     •    •     •    •     •    •     R1a1a1g1b   L260
•    •     •    •     •    •     •     R1a1a1g2   Z280
•    •     •    •     •    •     •    •     R1a1a1g2*   -
•    •     •    •     •    •     •    •     R1a1a1g2a    P278.2
•    •     •    •     •    •     •    •     R1a1a1g2b    L365
•    •     •    •     •    •     •    •     R1a1a1g2c    L366
•    •     •    •     •    •     •    •     R1a1a1g2d    Z92
•    •     •    •     •    •     •     R1a1a1g3   Z284
•    •     •    •     •    •     •    •     R1a1a1g3*   -
•    •     •    •     •    •     •    •     R1a1a1g3a   L448
•    •     •    •     •    •     •    •     •     R1a1a1g3a*   -
•    •     •    •     •    •     •    •     •     R1a1a1g3a1   L176.1/S179.1
•    •     •    •     •    •     •    •     •    •     R1a1a1g3a1*   -
•    •     •    •     •    •     •    •     •    •     R1a1a1g3a1a   L175
•    •     •    •     •    •     R1a1a1h   Z93
•    •     •    •     •    •     •     R1a1a1h*   -
•    •     •    •     •    •     •     R1a1a1h1   L342.2
•    •     •    •     •    •     •    •     R1a1a1h1*   -
•    •     •    •     •    •     •    •     R1a1a1h1a   L657
•    •     •    •     •    •     R1a1a1i   L664
•    •     R1b   M343
•    •     •     R1b*   -
•    •     •     R1b1   L278, M415, P25_1, P25_2, P25_3
•    •     •    •     R1b1*   -
•    •     •    •     R1b1a   P297, L320
•    •     •    •     •     R1b1a*   -
•    •     •    •     •     R1b1a1   M73, M478
•    •     •    •     •     R1b1a2   L265, M269, M520, S3, S10, S13, S17
•    •     •    •     •    •     R1b1a2*   -
•    •     •    •     •    •     R1b1a2a   L23/S141, L49.1
•    •     •    •     •    •     •     R1b1a2a*   -
•    •     •    •     •    •     •     R1b1a2a1   L150
•    •     •    •     •    •     •    •     R1b1a2a1*   -
•    •     •    •     •    •     •    •     R1b1a2a1a   L51/M412/S167
•    •     •    •     •    •     •    •     •     R1b1a2a1a*   -
•    •     •    •     •    •     •    •     •     R1b1a2a1a1   L11/S127, L52, L151, P310/S129, P311/S128
•    •     •    •     •    •     •    •     •    •     R1b1a2a1a1*   -
•    •     •    •     •    •     •    •     •    •     R1b1a2a1a1a   M405/S21/U106
•    •     •    •     •    •     •    •     •    •     •     R1b1a2a1a1a*   -
•    •     •    •     •    •     •    •     •    •     •     R1b1a2a1a1a1   P107
•    •     •    •     •    •     •    •     •    •     •     R1b1a2a1a1a2   L6
•    •     •    •     •    •     •    •     •    •     •     R1b1a2a1a1a3   L217
•    •     •    •     •    •     •    •     •    •     •     R1b1a2a1a1a4   Z18, Z19
•    •     •    •     •    •     •    •     •    •     •    •     R1b1a2a1a1a4*   -
•    •     •    •     •    •     •    •     •    •     •    •     R1b1a2a1a1a4a   L257/S186
•    •     •    •     •    •     •    •     •    •     •    •     R1b1a2a1a1a4b   L325
•    •     •    •     •    •     •    •     •    •     •     R1b1a2a1a1a5   Z381
•    •     •    •     •    •     •    •     •    •     •    •     R1b1a2a1a1a5*   -
•    •     •    •     •    •     •    •     •    •     •    •     R1b1a2a1a1a5a   Z156
•    •     •    •     •    •     •    •     •    •     •    •     •     R1b1a2a1a1a5a*   -
•    •     •    •     •    •     •    •     •    •     •    •     •     R1b1a2a1a1a5a1   L1/S26
•    •     •    •     •    •     •    •     •    •     •    •     •     R1b1a2a1a1a5a2   P89.2
•    •     •    •     •    •     •    •     •    •     •    •     R1b1a2a1a1a5b   Z301
•    •     •    •     •    •     •    •     •    •     •    •     •     R1b1a2a1a1a5b*   -
•    •     •    •     •    •     •    •     •    •     •    •     •     R1b1a2a1a1a5b1   M467/S29/U198
•    •     •    •     •    •     •    •     •    •     •    •     •     R1b1a2a1a1a5b2   L48/S162
•    •     •    •     •    •     •    •     •    •     •    •     •    •     R1b1a2a1a1a5b2*   -
•    •     •    •     •    •     •    •     •    •     •    •     •    •     R1b1a2a1a1a5b2a   L47/S170
•    •     •    •     •    •     •    •     •    •     •    •     •    •     •     R1b1a2a1a1a5b2a*   -
•    •     •    •     •    •     •    •     •    •     •    •     •    •     •     R1b1a2a1a1a5b2a1   L44/S171, L163
•    •     •    •     •    •     •    •     •    •     •    •     •    •     •    •     R1b1a2a1a1a5b2a1*   -
•    •     •    •     •    •     •    •     •    •     •    •     •    •     •    •     R1b1a2a1a1a5b2a1a   L46/S172
•    •     •    •     •    •     •    •     •    •     •    •     •    •     •    •     •     R1b1a2a1a1a5b2a1a*   -
•    •     •    •     •    •     •    •     •    •     •    •     •    •     •    •     •     R1b1a2a1a1a5b2a1a1   L45, L164, L237
•    •     •    •     •    •     •    •     •    •     •    •     •    •     R1b1a2a1a1a5b2b   Z9
•    •     •    •     •    •     •    •     •    •     •    •     •    •     •     R1b1a2a1a1a5b2b*   -
•    •     •    •     •    •     •    •     •    •     •    •     •    •     •     R1b1a2a1a1a5b2b1   Z2
•    •     •    •     •    •     •    •     •    •     •    •     •    •     •    •     R1b1a2a1a1a5b2b1*   -
•    •     •    •     •    •     •    •     •    •     •    •     •    •     •    •     R1b1a2a1a1a5b2b1a   Z7
•    •     •    •     •    •     •    •     •    •     •    •     •    •     •    •     •     R1b1a2a1a1a5b2b1a*   -
•    •     •    •     •    •     •    •     •    •     •    •     •    •     •    •     •     R1b1a2a1a1a5b2b1a1   Z8
•    •     •    •     •    •     •    •     •    •     •    •     •    •     •    •     •    •     R1b1a2a1a1a5b2b1a1*   -
•    •     •    •     •    •     •    •     •    •     •    •     •    •     •    •     •    •     R1b1a2a1a1a5b2b1a1a   Z11
•    •     •    •     •    •     •    •     •    •     •    •     •    •     •    •     •    •     •     R1b1a2a1a1a5b2b1a1a*   -
•    •     •    •     •    •     •    •     •    •     •    •     •    •     •    •     •    •     •     R1b1a2a1a1a5b2b1a1a1   Z12
•    •     •    •     •    •     •    •     •    •     •    •     •    •     •    •     •    •     •    •     R1b1a2a1a1a5b2b1a1a1*   -
•    •     •    •     •    •     •    •     •    •     •    •     •    •     •    •     •    •     •    •     R1b1a2a1a1a5b2b1a1a1a   L148/S173
•    •     •    •     •    •     •    •     •    •     •    •     •    •     •    •     •    •     R1b1a2a1a1a5b2b1a1b   Z1
•    •     •    •     •    •     •    •     •    •     •    •     •    •     •     R1b1a2a1a1a5b2b2   Z326
•    •     •    •     •    •     •    •     •    •     •    •     •    •     •    •     R1b1a2a1a1a5b2b2*   -
•    •     •    •     •    •     •    •     •    •     •    •     •    •     •    •     R1b1a2a1a1a5b2b2a   L188
•    •     •    •     •    •     •    •     •    •     •    •     •    •     R1b1a2a1a1a5b2c   L200
•    •     •    •     •    •     •    •     •    •     R1b1a2a1a1b   P312/S116
•    •     •    •     •    •     •    •     •    •     •     R1b1a2a1a1b*   -
•    •     •    •     •    •     •    •     •    •     •     R1b1a2a1a1b1   M65
•    •     •    •     •    •     •    •     •    •     •     R1b1a2a1a1b2   Z196
•    •     •    •     •    •     •    •     •    •     •    •     R1b1a2a1a1b2*   -
•    •     •    •     •    •     •    •     •    •     •    •     R1b1a2a1a1b2a   M153
•    •     •    •     •    •     •    •     •    •     •    •     R1b1a2a1a1b2b   L176.2/S179.2
•    •     •    •     •    •     •    •     •    •     •    •     •     R1b1a2a1a1b2b*   -
•    •     •    •     •    •     •    •     •    •     •    •     •     R1b1a2a1a1b2b1   M167/SRY2627
•    •     •    •     •    •     •    •     •    •     •    •     •     R1b1a2a1a1b2b2   L165/S68
•    •     •    •     •    •     •    •     •    •     •     R1b1a2a1a1b3   S28/U152
•    •     •    •     •    •     •    •     •    •     •    •     R1b1a2a1a1b3*   -
•    •     •    •     •    •     •    •     •    •     •    •     R1b1a2a1a1b3a   M126
•    •     •    •     •    •     •    •     •    •     •    •     R1b1a2a1a1b3b   M160
•    •     •    •     •    •     •    •     •    •     •    •     R1b1a2a1a1b3c   L2/S139
•    •     •    •     •    •     •    •     •    •     •    •     •     R1b1a2a1a1b3c*   -
•    •     •    •     •    •     •    •     •    •     •    •     •     R1b1a2a1a1b3c1   Z367
•    •     •    •     •    •     •    •     •    •     •    •     •    •     R1b1a2a1a1b3c1*   -
•    •     •    •     •    •     •    •     •    •     •    •     •    •     R1b1a2a1a1b3c1a   L20/S144
•    •     •    •     •    •     •    •     •    •     •    •     •    •     •     R1b1a2a1a1b3c1a*   -
•    •     •    •     •    •     •    •     •    •     •    •     •    •     •     R1b1a2a1a1b3c1a1   M228.2
•    •     •    •     •    •     •    •     •    •     •    •     •     R1b1a2a1a1b3c2   L196
•    •     •    •     •    •     •    •     •    •     •    •     •     R1b1a2a1a1b3c3   Z49
•    •     •    •     •    •     •    •     •    •     •    •     •    •     R1b1a2a1a1b3c3*   -
•    •     •    •     •    •     •    •     •    •     •    •     •    •     R1b1a2a1a1b3c3a   Z142
•    •     •    •     •    •     •    •     •    •     •    •     •    •     •     R1b1a2a1a1b3c3a*   -
•    •     •    •     •    •     •    •     •    •     •    •     •    •     •     R1b1a2a1a1b3c3a1   L562
•    •     •    •     •    •     •    •     •    •     •    •     R1b1a2a1a1b3d   Z36
•    •     •    •     •    •     •    •     •    •     •    •     R1b1a2a1a1b3e   Z56
•    •     •    •     •    •     •    •     •    •     •    •     •     R1b1a2a1a1b3e*   -
•    •     •    •     •    •     •    •     •    •     •    •     •     R1b1a2a1a1b3e1   L4/S178
•    •     •    •     •    •     •    •     •    •     •    •     •     R1b1a2a1a1b3e2   S47
•    •     •    •     •    •     •    •     •    •     •     R1b1a2a1a1b4   L21/M529/S145, L459
•    •     •    •     •    •     •    •     •    •     •    •     R1b1a2a1a1b4*   -
•    •     •    •     •    •     •    •     •    •     •    •     R1b1a2a1a1b4a   M37
•    •     •    •     •    •     •    •     •    •     •    •     R1b1a2a1a1b4b   M222/Page84/USP9Y+3636
•    •     •    •     •    •     •    •     •    •     •    •     R1b1a2a1a1b4c   DF1/L513/S215
•    •     •    •     •    •     •    •     •    •     •    •     •     R1b1a2a1a1b4c*   -
•    •     •    •     •    •     •    •     •    •     •    •     •     R1b1a2a1a1b4c1   P66_1, P66_2, P66_3
•    •     •    •     •    •     •    •     •    •     •    •     •     R1b1a2a1a1b4c2   L193/S176
•    •     •    •     •    •     •    •     •    •     •    •     R1b1a2a1a1b4d   L96
•    •     •    •     •    •     •    •     •    •     •    •     R1b1a2a1a1b4e   L144/S175, L195
•    •     •    •     •    •     •    •     •    •     •    •     R1b1a2a1a1b4f    L159.2/S169.2
•    •     •    •     •    •     •    •     •    •     •    •     R1b1a2a1a1b4g   Z253
•    •     •    •     •    •     •    •     •    •     •    •     •     R1b1a2a1a1b4g*   -
•    •     •    •     •    •     •    •     •    •     •    •     •     R1b1a2a1a1b4g1   L226/S168
•    •     •    •     •    •     •    •     •    •     •    •     •     R1b1a2a1a1b4g2    L554
•    •     •    •     •    •     •    •     •    •     •    •     R1b1a2a1a1b4h   DF21/S192
•    •     •    •     •    •     •    •     •    •     •    •     •     R1b1a2a1a1b4h*   -
•    •     •    •     •    •     •    •     •    •     •    •     •     R1b1a2a1a1b4h1   P314.2
•    •     •    •     •    •     •    •     •    •     •    •     •    •     R1b1a2a1a1b4h1*   -
•    •     •    •     •    •     •    •     •    •     •    •     •    •     R1b1a2a1a1b4h1a   L362
•    •     •    •     •    •     •    •     •    •     •    •     •     R1b1a2a1a1b4h2   Z246
•    •     •    •     •    •     •    •     •    •     •    •     •    •     R1b1a2a1a1b4h2*   -
•    •     •    •     •    •     •    •     •    •     •    •     •    •     R1b1a2a1a1b4h2a   DF25/S253
•    •     •    •     •    •     •    •     •    •     •    •     •    •     •     R1b1a2a1a1b4h2a*   -
•    •     •    •     •    •     •    •     •    •     •    •     •    •     •     R1b1a2a1a1b4h2a1   DF5/S191
•    •     •    •     •    •     •    •     •    •     •    •     •    •     •    •     R1b1a2a1a1b4h2a1*   -
•    •     •    •     •    •     •    •     •    •     •    •     •    •     •    •     R1b1a2a1a1b4h2a1a   L627
•    •     •    •     •    •     •    •     •    •     •    •     •     R1b1a2a1a1b4h3   L720
•    •     •    •     •    •     •    •     •    •     •    •     R1b1a2a1a1b4i    L371
•    •     •    •     •    •     •    •     •    •     •     R1b1a2a1a1b5   L238/S182
•    •     •    •     •    •     •    •     •    •     •     R1b1a2a1a1b6   DF19
•    •     •    •     •    •     •    •     R1b1a2a1b   L584
•    •     •    •     R1b1b   M335
•    •     •    •     R1b1c   V88
•    •     •    •     •     R1b1c*   -
•    •     •    •     •     R1b1c1   M18
•    •     •    •     •     R1b1c2   V8
•    •     •    •     •     R1b1c3   V35
•    •     •    •     •    •     R1b1c3*   -
•    •     •    •     •    •     R1b1c3a   V7
•    •     •    •     •     R1b1c4   V69
•     R2   M479
•    •     R2*   -
•    •     R2a   L266, M124, P249, P267
•    •     •     R2a*   -
•    •     •     R2a1   L295
•    •     •    •     R2a1*   -
•    •     •    •     R2a1a   L294
•    •     •     R2a2   L263

Link to Haplogroup R Private SNPs, Notes and References (Papers)

Y-DNA haplogroup R-M207 is believed to have arisen approximately 27,000 years ago in Asia. The two currently defined subclades are R1 and R2.

Link to Haplogroup R Private SNPs, Notes and References (Papers)

Additional Resources: (Projects arranged in the order that they appear on the tree.)

ISOGG Wiki - What you need to know about Genetic Genealogy.
Y-Haplogroup R* DNA Project, Vincent Vizacherro.
R-Arabia Y-DNA Project.
R1* Haplogroup Project, Vincent Vizacherro.
Y-DNA Haplogroup R1a - FTDNA vs. ISOGG Haplotrees, Diana Gale Matthiesen.
R1a International Y-DNA Project.
R1a1a and Subclades Y-DNA Project, Lukasz Lapinski, Lawrence Mayka, Marius Wilkoszewski, A. J. Levin, Anne M. Berge, Michal Milewski.
R1a WTY Project, Cyndi Rutledge.
Ashkenazi-Levite DNA Project (R1a1).
Y-DNA Haplogroup R1b — FTDNA vs. ISOGG Haplotrees, Diana Gale Matthiesen.
R1b and Subclades YDNA Haplogroup Project, Charles Kerchner.
The Jewish R1b Project, Elise Friedman, Sean Silver, Itzhak Epstein, Gil Bardige.
R1b1(xP297) DNA Project, Vincent Vizacherro, Peter Hrechdakian.
R1b1b1 (R1b1a1) Haplogroup Project (M73), Vincent Vizacherro.
R1b1a2 (P312- U106-) DNA Project (aka ht35 Project), Vincent Vizacherro, Peter Hrechdakian.
R1b1b2* (R1b1a2*) Project.
YDNA Haplogroup R1b-U106/S21+ Research Group, Dan Draghici.
R-U106xL48 WTY Project.
The U198 / S29 Project, John Sloan.
R-L48+ WTY Project, Charles Moore.
R1b-P89.2, Gary Blakely.
R-P312 and Subclades Project, Vincent Tilroe, Richard Stevens.
R-M153 The Basque Marker.
R1b - U152 & Subclades Research Project, Tibor Fehér, Steve Gilbert.
U152+/S28+ Data, David Faux.
Dal Riadic Migration Y Chromosome DNA Genealogy Page (L21), John McEwan.
R-L21 Plus Project, Vincent Tilroe, Richard Stevens,
R-DF21 and Subclades, David Reynolds.
R-L21 Walk Through the Y, Kirsten Saxe, Vincent Tilroe, Alex Williamson.
R1b DYS464X=15.3g Subclade, David Reynolds.
R-L21* 11-13 Combo: includes L193+, P314.2+, Adam Bradford.
R-M222 Haplogroup Project (Niall of the 9 Hostages, Ui Neill Clan), David Wilson.
R-L159 Project.
The R-L226 Project - Irish Type III, Dennis O'Brien, Dennis Wright.
The Irish Type III Website (L226) (Dalcassian Clans), Dennis Wright.
The Y-Haplogroup R SRY2627+ and L176.2+ Project, Gareth Henson.
R-L165 (S68) Project, Lori McLeod-Wilke, Alasdair Macdonald, Conrad Terrill, Timothy McLeod.
R2 Y-Chromosome Haplogroup DNA Project.
R2-M124-WTY Project.

Corrections/Additions made since 1 January 2011.

Contact people for Haplogroup R:
R1a Contact: Lawrence Mayka
R-U106 and Subclades Contact: Charles Moore
R-P312 and Subclades Contact: Vince Tilroe Remainder of R Tree Contact: David Reynolds

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